B-4
<br />
<br />lodges and bank dens, depending
<br />on the situation. All lodges and
<br />bank dens have at least two en-
<br />trances and may have four or more.
<br />The ledge or bank den is used
<br />primarily for raising young, sleep-
<br />ing and some food storage during
<br />severe weather (Figure 5).
<br />
<br />The length or height of a dam
<br />generally depends upon what is
<br />necessary to essentially stop the
<br />flow of water and create a pond.
<br />In areas of flat topography, the
<br />dam, re, ay, not be~,.ov~,~. 3~, i. nches,.(.9~,.
<br />m) high, but as much as 1/4 mile (.4
<br />km) long. In hilly or mountain
<br />country, the dam may be q0 feet (3
<br />m) high and only 50 feet (15 m)
<br />long. The size of the tree the
<br />beaver cuts is highly variable, from
<br />a 1-inch (2.54 cm) DBH (diameter
<br />at breast height) softwood to a six-
<br />foot (1.8 m) DBH hardwood. In
<br />some areas beavers seem to cut
<br />down trees up to about 10 inches
<br />(25 cm) DBH and merely girdle or
<br />partially cut larger ones, although
<br />they often cut down much larger
<br />trees. For some reason the bea~ers
<br />seem to like to girdle large pines
<br />and sweetgums. They like the gum
<br />or storax that seeps out of the
<br />girdled area of sweetgum and
<br />other species.
<br />
<br />An important factor about beaver
<br />biology is thei~ territoriality. A col-
<br />ony generally consists of four to
<br />eight related beavers, who resist
<br />additions or outsiders to the col-
<br />ony or the pond. Young beavers
<br />commonly are displaced from the
<br />colony shortly after they become
<br />sexually mature, at about 2 years
<br />old. They often move to another
<br />area to begin a new pond and col-
<br />ony. However, some become
<br />solitary hermits inhabiting old
<br />abandoned ponds or a farm pond
<br />if one is available.
<br />
<br />Beavers have a few natural
<br />predators aside from man, in-
<br />cluding coyote, bobcat, river otter
<br />and mink on young kittens. In
<br />other areas, bear, wolves, and
<br />wolverines may prey on beavers.
<br />They are hosts for several ectopar-
<br />
<br />Figure 6. Pine plantation in Arkansas killed
<br />by floodin8 caused by beavers.
<br />
<br />asites and internal parasites in-
<br />cluding nematodes, trematodes
<br />and coccidians. Giardia lamblia is a
<br />pathogenic intestinal parasite
<br />transmitted by beavers, which has
<br />caused human health problems in
<br />water supply systems. In fact, the
<br />Centers for Disease Control have
<br />reports of at least 41 waterborne
<br />Giardiasis outbreaks affecting more
<br />than 15,000 people. For more infor-
<br />mation about Giardiasis, see von
<br />Oettingen (1982). 'This thesis con-
<br />tains a number of additional
<br />references.
<br />
<br />They generally are monogamous;
<br />copulation may take place either
<br />in 'the. water or in the lodge or
<br />bank den. They communicate by
<br />vocalizations, posture, tail slapping
<br />and scent posts or mud dobs or
<br />mounds placed around the bank
<br />and dam. The beaver's castor
<br />glands secrete a substance that is
<br />deposited on mud dobs to mark
<br />territorial boundaries. These scent
<br />posts are found more frequently at
<br />certain seasOns, but.are found
<br />year-round in active ponds.
<br />The habitat modification by
<br />beavers, caused primarily by dam
<br />building, is Often beneficial to fish,
<br />furbearers, reptiles and amphibians
<br />
<br />and to waterfowl and shorebirds.
<br />However, when this modification
<br />comes in conflict with man's objec-
<br />tives, the impacts of damage may
<br />far outweigh the benefits.
<br />
<br />Damage and Damage
<br />Identification
<br />
<br />Most of the damage caused by
<br />beavers is a result of dam building,
<br />bank burrowing, tree cutting or
<br />flooding. Some individual south-
<br />eastern states where beaver~dam-
<br />age is extensive have determined
<br />from surveys that beaver damage
<br />estimates range from $3 million to
<br />$5 million dollars annually to
<br />timber loss, crop losses, roads,
<br />dwel in&~,, a.nd property flooded,
<br />'and other damage. In some of
<br />these states, bottomland hardwood
<br />timber in areas of several thousand
<br />acres in one watershed maybe lost
<br />because of beaVer pond flooding.
<br />Some unusual cases observed over
<br />the years include state highways
<br />flooded because of beaver ponds,
<br />reservoir dams destroyed by bank
<br />den burrows collapsing, and train
<br />derailments caused by burrowing.
<br />Housing developments have been
<br />threatened by beaver dam flood-
<br />ing, and thousands of acres of
<br />croplands and young pine planta-
<br />
<br />
<br />
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